Model averaging real parameters

Hi folks,
I apologize if these questions don't make sense due to a fundamental (statistical) misunderstanding on my part....
1) I understand that in RMark model averaging is over the real parameters and not the betas (due to additional complications involved with averaging the betas - good discussion here: viewtopic.php?f=1&t=996&p=2620&hilit=model+average+betas#p2620). The help for model.average.list states that RMark uses eqtn 4.1 from B&A. What I cannot tell from the eqtn is how averaging is done when a predictor is not in every model (same as the beta discussion in the above link): are real parameters averaged only over the models that contain a given predictor (the "natural" average) or they are averaged over the entire model set, where a given real parameter is considered zero for a model that does not contain that predictor (the "zero" method)? Or does this question only make sense in the context of model averaging betas?
2) Assume a model set where some models contain interactions and other models contain the same predictors but without an interaction. If this set is model averaged, can one interpret the 'main effects' of the predictors? Grueber et al. 2011 (J. of Evo Bio) discuss the importance of 'centralizing' predictors when model sets have interactions. I understand that MARK standardizes covariates internally but I'm not sure whether that's relevant for this example.
Thanks!
Joe
I apologize if these questions don't make sense due to a fundamental (statistical) misunderstanding on my part....

1) I understand that in RMark model averaging is over the real parameters and not the betas (due to additional complications involved with averaging the betas - good discussion here: viewtopic.php?f=1&t=996&p=2620&hilit=model+average+betas#p2620). The help for model.average.list states that RMark uses eqtn 4.1 from B&A. What I cannot tell from the eqtn is how averaging is done when a predictor is not in every model (same as the beta discussion in the above link): are real parameters averaged only over the models that contain a given predictor (the "natural" average) or they are averaged over the entire model set, where a given real parameter is considered zero for a model that does not contain that predictor (the "zero" method)? Or does this question only make sense in the context of model averaging betas?
2) Assume a model set where some models contain interactions and other models contain the same predictors but without an interaction. If this set is model averaged, can one interpret the 'main effects' of the predictors? Grueber et al. 2011 (J. of Evo Bio) discuss the importance of 'centralizing' predictors when model sets have interactions. I understand that MARK standardizes covariates internally but I'm not sure whether that's relevant for this example.
Thanks!
Joe